Characterization of Intersubunit Communication in the Virginiamycin trans-Acyl Transferase Polyketide Synthase.

Fiche publication


Date publication

mars 2016

Journal

Journal of the American Chemical Society

Auteurs

Membres identifiés du Cancéropôle Est :
Dr GRUEZ Arnaud, Dr JACOB Christophe, Pr WEISSMAN Kira


Tous les auteurs :
Dorival J, Annaval T, Risser F, Collin S, Roblin P, Jacob C, Gruez A, Chagot B, Weissman KJ

Résumé

Modular polyketide synthases (PKSs) direct the biosynthesis of clinically valuable secondary metabolites in bacteria. The fidelity of chain growth depends on specific recognition between successive subunits in each assembly line: interactions mediated by C- and N-terminal "docking domains" (DDs). We have identified a new family of DDs in trans-acyl transferase PKSs, exemplified by a matched pair from the virginiamycin (Vir) system. In the absence of C-terminal partner (VirA (C)DD) or a downstream catalytic domain, the N-terminal DD (VirFG (N)DD) exhibits multiple characteristics of an intrinsically disordered protein. Fusion of the two docking domains results in a stable fold for VirFG (N)DD and an overall protein-protein complex of unique topology whose structure we support by site-directed mutagenesis. Furthermore, using small-angle X-ray scattering (SAXS), the positions of the flanking acyl carrier protein and ketosynthase domains have been identified, allowing modeling of the complete intersubunit interface.

Mots clés

Acyltransferases, metabolism, Magnetic Resonance Spectroscopy, Models, Molecular, Polyketide Synthases, metabolism, Virginiamycin, chemistry

Référence

J. Am. Chem. Soc.. 2016 Mar;138(12):4155-67